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Information Plants usually on rock. Stems compact to long-creeping, ascending to horizontal, usually branched; scales brown to tan or often bicolored with dark, central stripe and lighter margins, linear-subulate to lanceolate (rarely ovate), margins dentate, erose, or entire. Leaves monomorphic to somewhat dimorphic, clustered to widely scattered, 2--100 cm. Petiole brown, black, straw-colored, or gray, rounded, flattened or with single longitudinal groove adaxially, glabrous or pubescent, usually with a few scales at base, with single vascular bundle. Blade linear to ovate-deltate, 1--4-pinnate proximally, leathery or rarely somewhat herbaceous, abaxially glabrous, pubescent, or with hairlike scales scattered along costae, adaxially usually glabrous, dull, not striate; rachis straight or flexuous. Ultimate segments of blade usually stalked and free from costae, elliptic, lanceolate to linear, usually more than 4 mm wide; base rounded, truncate, or cordate; stalks often lustrous and dark colored; segment margins reflexed to form confluent, poorly defined, false indusia extending entire length of segment. Veins of ultimate segments free or rarely anastomosing, usually obscure, pinnately branched and divergent distally. False indusia greenish to whitish, narrow, clearly marginal, often concealing the sporangia. Sporangia scattered along veins near segment margins, containing 32 or 64 spores, often intermixed with glands, farina-producing. Spores brown to tan (rarely yellow), tetrahedral-globose, rugose or cristate, lacking prominent equatorial ridge. x = 29.

Information Stems compact, ascending, stout, 5--10 mm diam.; scales bicolored, linear-subulate, 0.1--0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate. Leaves somewhat dimorphic, sterile leaves shorter and less divided than fertile leaves, clustered on stems, 8--40 cm; croziers sparsely villous. Petiole dark brown, lustrous, flattened or slightly grooved adaxially, without prominent articulation lines. Blade ovate-deltate, usually 2-pinnate proximally, 4--18 cm wide; rachis brown throughout, straight, shallowly grooved adaxially, usually glabrous. Pinnae perpendicular to rachis to slightly ascending, not decurrent on rachis, usually with 9--25 ultimate segments; costae straight, 20--70 mm, much longer than fertile ultimate segments. Ultimate segments narrowly oblong, 4--10 mm, leathery, glabrous; margins recurved on fertile segments, usually covering less than 1/2 abaxial surface, borders whitish, nearly entire; apex mucronate. Veins of ultimate segments obscure. Sporangia long-stalked, containing 64 spores, intermixed with abundant farina-producing glands. 2 n = 58.

Information Stems compact, ascending, stout, 5--10 mm diam.; scales bicolored, linear-subulate, 0.1--0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate. Leaves monomorphic, clustered on stem, 6--40 cm; croziers sparsely villous. Petiole dark brown, lustrous, flattened or slightly grooved adaxially, without prominent articulation lines. Blade linear-oblong, 2-pinnate proximally, 1.5--5 cm wide; rachis brown throughout, straight, shallowly grooved adaxially, usually glabrous. Pinnae perpendicular to rachis or slightly ascending, not decurrent on rachis, usually with 3--9 ultimate segments; costae straight, 2--20 mm, usually shorter than ultimate segments. Ultimate segments narrowly oblong, 5--20 mm, leathery, glabrous; margins recurved on fertile segments, usually covering less than 1/2 abaxial surface, borders whitish, crenulate; apex mucronate. Veins of ultimate segments obscure. Sporangia long-stalked, containing 64 spores, intermixed with sparse farina-producing glands. 2 n = 116.

Information Stems compact, ascending, stout, 5--10 mm diam.; scales uniformly brown or tan, linear-subulate, 0.1--0.3 mm wide, thin, margins entire to denticulate. Leaves somewhat dimorphic, sterile leaves shorter and less divided than fertile leaves, clustered on stem, 10--30 cm; croziers villous. Petiole dark brown to reddish purple, lustrous, rounded adaxially, without prominent articulation lines. Blade elongate-deltate to ovate, 2-pinnate proximally, 5--15 cm wide; rachis brown or reddish purple throughout, straight, often slightly flattened adaxially, sparsely villous with long, divergent hairs. Pinnae perpendicular to rachis or slightly ascending, not decurrent on rachis, usually with 7--15 ultimate segments; costae straight, 25--80 mm, usually longer than ultimate segments. Ultimate segments oblong-lanceolate, 7--25 mm, leathery, sparsely villous abaxially near midrib; margins usually recurved on fertile segments, covering less than 1/2 abaxial surface, borders whitish, crenulate; apex slightly mucronate. Veins of ultimate segments obscure. Sporangia long-stalked, containing 32 spores, not intermixed with farina-producing glands.

Information Stems compact, ascending, stout, 5--10 mm diam.; scales bicolored, linear-subulate, 0.1--0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate. Leaves monomorphic, clustered on stem, 7--45 cm; croziers sparsely villous. Petiole dark brown, lustrous, flattened to slightly grooved adaxially, without prominent articulation lines. Blade ovate-deltate, (2--)3-pinnate proximally, 4--18 cm wide; rachis brown throughout, straight, shallowly grooved adaxially, usually glabrous. Pinnae perpendicular to rachis to strongly ascending, not decurrent on rachis, usually with 9--40 ultimate segments; costae straight, 10--70 mm, much longer than ultimate segments. Ultimate segments narrowly oblong, 2--12 mm, leathery, glabrous; margins recurved to strongly revolute on fertile segments, usually covering more than 1/2 abaxial surface, borders greenish, usually dentate; apex mucronate. Veins of ultimate segments obscure. Sporangia short-stalked, containing 64 spores, intermixed with abundant farina-producing glands.

Information Stems compact, ascending, stout, 5--10 mm diam.; scales uniformly reddish brown (or tan), linear-subulate, 0.1--0.3 mm wide, thin, margins entire to denticulate. Leaves somewhat dimorphic, sterile leaves shorter and less divided than fertile leaves, clustered on stems, 5--50 cm; croziers villous. Petiole reddish purple to nearly black, lustrous, rounded adaxially, without prominent articulation lines. Blade elongate-deltate, usually 2-pinnate proximally, 2--18 cm wide; rachis reddish purple throughout, straight, rounded adaxially, densely pubescent adaxially with short, curly, appressed hairs. Pinnae perpendicular to rachis or ascending, not decurrent on rachis, usually with 3--15 ultimate segments; costae straight, 10--100 mm, often longer than ultimate segments. Ultimate segments linear-oblong, 10--75 mm, leathery, sparsely villous abaxially near midrib; margins weakly recurved to plane on fertile segments, usually covering less than 1/2 abaxial surface, borders whitish, crenulate; apex obtuse to slightly mucronate. Veins of ultimate segments obscure. Sporangia long-stalked, containing 32 spores, not intermixed with farina-producing glands. n = 2 n = 87, apogamous.